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Since Spiderfang units move over terrain as if they can fly, are they able to shoot through the Wyldwoods? Or do they have to be able to outright fly? In which case they'd be BeeFangs instead

Not normally because "do X as if they can fly" is not the same as "can fly."

However I believe there was an errata somewhere that said if you sculpt wings on all your spiders and make little buzzing noises when moving them they do indeed gain the full fly ability.

I’m totally doing that, and I’m gonna dance the next time I use the Masque too!