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Is there a rule that prohibits the trees from being removed from the base and placing units in the spot where the tree was previously? I have played it this way for a while, but a friend recently told me there was a GW battle where they played it as though the trees were removed and models could be placed anywhere on the wyldwood base.

The trees have always been removable so you could position units in the forest as needed. You're able to remove the trees. As long as they're in the forest, they're in cover. The forest is the base.

I remember reading an article from the designers that they were not supposed to be removed. trees should be climbed and considered as any other scenery. There is no rule that says they should be treated otherwise., and if a model cannot pass because the base is to big, it cannot pass.

In tournaments I have attended people can remove the trees, but are not allowed to put bases where the trees were... (I guess to make sure that branches do not stop a treelord from entering a forest....)

 minisnatcher wrote:
I remember reading an article from the designers that they were not supposed to be removed. trees should be climbed and considered as any other scenery. There is no rule that says they should be treated otherwise., and if a model cannot pass because the base is to big, it cannot pass.

In tournaments I have attended people can remove the trees, but are not allowed to put bases where the trees were... (I guess to make sure that branches do not stop a treelord from entering a forest....)

So apparently in the March Edition of White dwarf, there are pictures of a battle report where a model is placed over the holes where the trees belong on a wyldwood. This is why I am asking for something documented regarding the rules at large. Without something definitive, its just conjecture one way or the other.

The 'definitive' line you're looking for is the rule that models cannot move through solid terrain models unless explicitly allowed to do so (so your infantry can't walk through walls, etc), and the fact that there is no rule anywhere saying you can remove trees from the wyldwoods base.

The practice of leaving the wyldwoods trees unglued and removing them for ease of access is a strictly 'I don't want to send my models flying because a branch snagged on my sleeve' thing, there's no rule that actually allows you to not finish assembling the model and utilize the amount of space that frees up, so in absence of the tree you should play as iff the hole is not passible (though to be fair plenty of models could easily sit on top of those roots).

The only time this came up for me was in a game where a guy tried to charge in with his zombie dragon and we found he couldn't actually fit it anywhere inside the forest. Of course we then realized since it could fly it could sit on top of the trees for maximum sillyness.

I suppose removing the trees and stating the model is on top of them is one way to play it, as AoS lacks the 40k rule of "models cannot be placed on top of other models". Otherwise it seems a tad silly that a Branchwych in the center of a forest is immune to a charge from a creature like a zombie dragon type of model.

If the model is on top of the trees, it will most likele not be able to hit anything on the ground level as all distances are measured base to base. so a charge would never finish within 0,5 inch and I think they are certainly more then 2" high so most attacks would iss. Wobbly model can only be used if the player can balance it that way and then remove it because it would otherwise fall. So a branchwych in the middle of a forest is possibly immune for a zombie dragon charge depending on how the trees and models are placed.