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I love this game but I am still trying to figure out some of the sequencing. If you take Bossk with a Mangler Can you convert a hit to a crit with the MC and then convert the crit to two hits with Bossk's ability?

Yep.

You may only reroll a dice once, but you may modify them multiple times.

In a similar way you may score a crit with a Heavy Laser Cannon by modifying the crits to hits, then using another effect to convert them back.

Yes, but not the way it sounds like you want.

Modifying the dice multiple times is fine, but Bossk's ability only triggers on an uncanceled crit. You roll dice for your attack, the defender modifies attack dice (if able), you modify your attack dice, the defender rolls green dice, you modify their defense dice (if able), they modify their defense dice, then you compare results. If, after you compare results, there is an uncanceled crit, you may cancel that result and add two hits. The thing to remember is that there are two distinct events happening here: modifying and canceling.

Thanks SPJ. I reread the attack section of the rule book and now I get it, because it says you apply Bossks ability before assigning hits it is happening between steps 6 and 7 in the rule book.

As an ESL speaker, I love that this game is so clearly defined, the rules only require that you actually read them, then if you have questions, it is just down to reading comprehension, not interpretive theology, with a healthy dose of comparative anthropology.