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Do skaven get their rank bonus for leadership tests while in buildings? They don't get their ranks in combat, but I don't recall other restrictions (and, from a fluff perspective, they should probably recognize that all their friends are still around them (perhaps it's a building with lots of really small rooms?)

There are no rank bonuses in building assaults, for either side.
So no streinght in numbers for the poor rats.

The two dont necessarily follow - just because your ranks dont count for determining combat res doesnt mean they necessarily lose their ranks in toto.

Nothing in the rules, that I could see, state that a unit loses its ranks when it enters a building. The only time ranks are even mentioned (apart from leaving the building) is in assault results.

So unless there is something written differently you still have your ranks while within the building, so SiN would still apply.

That does sound reasonable.