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what happens is sorta this.

marine placed into 10th company as scout.

serves for so many years or for so many actions until his sergeant puts him up for promotion. This can be sped up if battle companies need replacements.

He then gets placed in either the devestators or assault marine battle companies for x amount of time and then swaps to the other for another x amount of time. He is then promoted to an active battle company (3rd-7th). Other options at this point are recruitment into the Apothecarion, Libraius, or Mechanicum in which they leave the chapter for a time to train for their new role.

Marines are stuck here until they are judged competent enough to be promoted. They will either take up a sergeant position in one of the battle companies, or (in the case of a wounded or disgraced marine) possibly be sent back home to lead new recruits in the 10th or be stationed with the 2nd company defence force.

If a marine serves long enough as a serg in an active battle company he can gain veteran status, at which point he will either be promoted to first company or take up a position in his current battle company's command squad.

Truly heroic and capable marines who serve dilligently in the first company may, upon the death of a previous captain, receive a promotion to captain of one of the companies.