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Guilliman was the most human.

This is a problem, since his bros were largely not-human. He got along with pretty much all the sensible military and more human brothers. It makes sense that he was best buds with Dorn, admired Sanguinius (like everyone else), had respect for the Lion, who was an intellectual and confident leader, Ferrus who was a fair but demanding guy, while having respect for the Khan who handled being a warrior with dignity.

The others?
Horus was a show-off, but Guilliman respected his accomplishments.
Fulgrim was an fop with obvious issues.
Lorgar and Magnus were mystics, recluses and weirdos.
Conrad was a murdering ass.
Angron was a murdering ass.
Russ is an animal who keeps trying to be a big man. The Khan was at least reserved.
Mortarion was a distant, cold jerk.
Alpharius always was trying to be clever and mysterious for it's own sake.
Perturabo was a brooding ass prone to stupid rages.
Corax and Vulkan just did their own thing.

Sanguinius was not 'furthest from falling'. He was almost sort of guilted into it, with all of that red angel nonsense.

One issue is that Guilliman was more successful than even Horus in a number of ways. Horus may have conquered more worlds, but Guilliman made more productive worlds, with zero internal unrest, and huge industrial/manpower outputs.

He also had the most marines, trained to the highest standards (much better per-soldier it's suggested than the iron warriors and word bearers for example).

His human modesty is a lot more normal than the Lion's brooding distance for example. His methods worked, and were building a much better Imperium than whatever Curze, Alpharius and Perturabo were doing.

He didn't really seem to have beef with Russ, Khan or Magnus, who while he may not have liked, weren't really messing things up. They were doing a specific purpose, like hunting down particularly nasty enemies. But Curze and Angron were definitely just making things worse.

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Horus getting along with everyone was because he was flashy. He was an adventurer, who was personally very involved with war, and neurotic about post-war lack of glory.

Guilliman was a statesman, who was happy to build the best planets and imperium possible, and knew that doing stuff like that would always land a job. So he didn't really care so much about glory, or even seeking the approval of his crazier brothers. He's also probably awkward because unlike everyone else, he had a /Mom/, and a Dad who was actually a really good guy who he looked up to, before Empy showed up. (Almost) everyone else either had no parents but a tribe/group raising them, animal parents, or evil parents who had to die.