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So in the Death Guard Vectorum all units that can must take Mark of Nurgle. I'm wanting to take Typhus as my warlord. His Plague Zombie rule states that only non-upgraded units can be turned to zombies. Does this mean I can't use his rule at all, since my cultists are forced to take Mark of Nurgle?

The change to zombies is made before they can take any options. Therefore by the time you get to the Mark option, the unit is banned from taking it.