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When Horticulous Slimux plants a tree with his special rule, does it cost reinforcement points?

Yes, it does. More generally, replacing slain models to a unit (by default) does not cause reinforcement points, adding new models above its starting strength does cost reinforcement points and creating new units (such as adding a new Feculent Gnarlmaw, for example) costs reinforcement points. This can, of course, be overridden by a units own specific rules (such as the case for Horrors).

Is there a rule you can point to for that?

 Jidmah wrote:
Is there a rule you can point to for that?

Yes, the Matched Play Reinforcement Points rules.

BRB page 214 wrote:Each time a unit is added to an army during battle, you must first subtract the number of points the unit would cost from your pool of reinforcement points.

The Seed the Garden of Nurgle rule states "add a new Feculent Gnarlmaw (pg 118) to your army".

Perfectly clear, thanks