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-DA Chapter Ancient: DARK ANGELS INFANTRY models that are destroyed within 6" of any friendly DARK ANGELS CHAPTER ANCIENTS and subsequently muster one last surge of strength before succumbing to their wounds (see Astartes Banner ability, above) always resolve their final attack as if they had a Ballistic Skill and a Weapon Skill characteristic of 2+.

-Culexus Ability: Each time an enemy model attacks this model, the attacks are resolved as if the the attacker had a BS/WS of 6+

If the DA unit is shooting at the Assassin, which takes precedence?

Q: If a model whose attacks always hit on a specific roll (such as
a Vindicare Assassin or Dark Reaper’s ranged attacks) targets
a unit that can only be hit on a specific roll (such as a unit
under the effects of the Pathfinders Stratagem), which rule takes
precedent – the attacking model’s or the target unit’s?
A: The attacking model’s ability takes precedence.

 p5freak wrote:

Q: If a model whose attacks always hit on a specific roll (such as
a Vindicare Assassin or Dark Reaper’s ranged attacks) targets
a unit that can only be hit on a specific roll (such as a unit
under the effects of the Pathfinders Stratagem), which rule takes
precedent – the attacking model’s or the target unit’s?
A: The attacking model’s ability takes precedence.

Neither of these are "always hits on a X+" abilities.
They are both "the attacks are resolved as if the attack has a BS/WS of X+"
It's different syntax and that FAQ does not apply.

Unfortunately, I can not seem to find the answer... I would say the closest is thing for precedence is this quoted FAQ but its a house rule because no RAW seems to exist that I can find.

PapaHorror wrote:
-DA Chapter Ancient: DARK ANGELS INFANTRY models that are destroyed within 6" of any friendly DARK ANGELS CHAPTER ANCIENTS and subsequently muster one last surge of strength before succumbing to their wounds (see Astartes Banner ability, above) always resolve their final attack as if they had a Ballistic Skill and a Weapon Skill characteristic of 2+.

-Culexus Ability: Each time an enemy model attacks this model, the attacks are resolved as if the the attacker had a BS/WS of 6+

If the DA unit is shooting at the Assassin, which takes precedence?

Neither do, because GW can't write rules. You have to invoke TMIR and roll off.

I have always seen it played as attacking unit takes precedence in situations like this.

DA takes precedence since it says always.