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The smaller Squiggoth might work. A Gorkanaut (or possibly Morkanaut?) would be also work. It's a little bit smaller than those, but with the Megaboss on top, you could convert him to be carrying a a large weapon to count for its guns.
Really, the Maw-crusha is big beefy wyvern, and can fly across the battlefield. So is there something that you could use with that in mind?