It depends on the line of sight of each model, if that model can see any part of the target then it can shoot/cast at it. Generally this means you can shoot through anything short of a building or other totally solid object, since being able to see the tip ofone enemy's oversized mohawk is enough to target him and his whole unit. Also, the rules call out that you can ignore other models from the same unit for the purposes of line of sight.
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